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I cannot replicate this bug, but the problem seems to be in the empty subSubsystems entries. A workaround (only works for the tutorial):

protRxns = contains(ecModel.rxns,'prot_');
ecModel.subSystems(protRxns) = ecModel.subSystems(protRxns) = {{'Protein usage reaction'}};

I will change that makeEcModel defines non-empty subsystems for the protein usage reactions.

This also highlights a bug in RAVEN's readYAMLmodel, which loads subSystems as nested cell arrays, while it should be "just" a cell array.

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@edkerk
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Answer selected by vivarock233-byte
@edkerk
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@vivarock233-byte
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