diff --git a/src/betterproto/__init__.py b/src/betterproto/__init__.py
index 82a5b1489..5a586c924 100644
--- a/src/betterproto/__init__.py
+++ b/src/betterproto/__init__.py
@@ -1344,6 +1344,8 @@ def to_pydict(
         defaults = self._betterproto.default_gen
         for field_name, meta in self._betterproto.meta_by_field_name.items():
             field_is_repeated = defaults[field_name] is list
+            if self.__raw_get(field_name) is PLACEHOLDER and not include_default_values:
+                continue
             value = getattr(self, field_name)
             cased_name = casing(field_name).rstrip("_")  # type: ignore
             if meta.proto_type == TYPE_MESSAGE:
@@ -1388,13 +1390,7 @@ def to_pydict(
 
                 if value or include_default_values:
                     output[cased_name] = value
-            elif (
-                value != self._get_field_default(field_name)
-                or include_default_values
-                or self._include_default_value_for_oneof(
-                    field_name=field_name, meta=meta
-                )
-            ):
+            else:
                 output[cased_name] = value
         return output
 
@@ -1476,7 +1472,6 @@ def _validate_field_groups(cls, values):
         field_name_to_meta = cls._betterproto_meta.meta_by_field_name  # type: ignore
 
         for group, field_set in group_to_one_ofs.items():
-
             if len(field_set) == 1:
                 (field,) = field_set
                 field_name = field.name
diff --git a/src/betterproto/grpc/grpclib_server.py b/src/betterproto/grpc/grpclib_server.py
index 5c1f93452..3e2803113 100644
--- a/src/betterproto/grpc/grpclib_server.py
+++ b/src/betterproto/grpc/grpclib_server.py
@@ -21,7 +21,6 @@ async def _call_rpc_handler_server_stream(
         stream: grpclib.server.Stream,
         request: Any,
     ) -> None:
-
         response_iter = handler(request)
         # check if response is actually an AsyncIterator
         # this might be false if the method just returns without
diff --git a/src/betterproto/plugin/compiler.py b/src/betterproto/plugin/compiler.py
index 7542432b2..510d64857 100644
--- a/src/betterproto/plugin/compiler.py
+++ b/src/betterproto/plugin/compiler.py
@@ -21,7 +21,6 @@
 
 
 def outputfile_compiler(output_file: OutputTemplate) -> str:
-
     templates_folder = os.path.abspath(
         os.path.join(os.path.dirname(__file__), "..", "templates")
     )
diff --git a/src/betterproto/plugin/parser.py b/src/betterproto/plugin/parser.py
index 358cc20e4..f48533338 100644
--- a/src/betterproto/plugin/parser.py
+++ b/src/betterproto/plugin/parser.py
@@ -159,7 +159,6 @@ def _make_one_of_field_compiler(
     proto_obj: "FieldDescriptorProto",
     path: List[int],
 ) -> FieldCompiler:
-
     pydantic = output_package.pydantic_dataclasses
     Cls = PydanticOneOfFieldCompiler if pydantic else OneOfFieldCompiler
     return Cls(
diff --git a/tests/inputs/google_impl_behavior_equivalence/test_google_impl_behavior_equivalence.py b/tests/inputs/google_impl_behavior_equivalence/test_google_impl_behavior_equivalence.py
index 0297692c6..b51851a3d 100644
--- a/tests/inputs/google_impl_behavior_equivalence/test_google_impl_behavior_equivalence.py
+++ b/tests/inputs/google_impl_behavior_equivalence/test_google_impl_behavior_equivalence.py
@@ -13,7 +13,6 @@
 
 
 def test_oneof_serializes_similar_to_google_oneof():
-
     tests = [
         (Test(string="abc"), ReferenceTest(string="abc")),
         (Test(integer=2), ReferenceTest(integer=2)),
@@ -30,7 +29,6 @@ def test_oneof_serializes_similar_to_google_oneof():
 
 
 def test_bytes_are_the_same_for_oneof():
-
     message = Test(string="")
     message_reference = ReferenceTest(string="")